Skip to main content
The University of Chicago Marine Biological Laboratory
expanded
collapsed
Weather
  • VISIT THE MBL
Utility
  • Contact
  • Careers
  • Give
    • About the MBL
    • Leadership
      • Leadership - Overview
      • Board of Trustees Committees
      • Nipam Patel, Director
        • Nipam Patel, Director - Overview
        • Gallery of Nipam Patel Research Images
      • Courtney Buckley
      • Alison Crawford
      • Samantha Cummis
      • Mary S. Harrington
      • Linda Hyman
      • Scott Koerner
      • Marie Russell
      • Sharon Stanczak
      • Anne W. Sylvester
      • Jennifer Walton
      • MBL Director History
    • Advisory Bodies
      • Advisory Bodies - Overview
        • MBL Society - Overview
        • Anne Hart
        • MBL Society Members
        • MBL Society Membership Criteria and Procedures
        • MBL Community Advisory Council Charter
        • MBL Science Council Charter
        • MBL Society Steering Committee
    • MBL Community
      • MBL Community - Overview
      • Land Acknowledgement Statement
      • MBL Community Committee
    • University of Chicago
    • Financial Statements
    • Careers & Internships
      • Careers & Internships - Overview
      • Our Community
      • Benefits Summary
        • Equal Employment Opportunity - Overview
        • Accommodation of Individuals with Disabilities
        • Drug-Free Workplace
        • Harassment-Free Workplace
      • Notice of Nondiscrimination
      • Staff Toolbox
      • Electronic Posters
      • Contact HR
    • History & Archives
      • History & Archives - Overview
      • Nobel Laureates Affiliated with the MBL
        • Legacy of Leadership - Overview
        • Cornelia Clapp
        • Jewel Plummer Cobb
        • Jean Clark Dan and Katsuma Dan
        • Ernest Everett Just
        • Stephen W. Kuffler
        • Joseph L. Martinez, Jr. and James G. Townsel
        • Lilian Vaughan and Thomas Hunt Morgan
        • Seeing Life Exhibit - Overview
        • Seeing Life at MBL
        • Bronze Memorial Tablet
        • Charles Otis Whitman
        • Edmund Beecher Wilson
        • Thomas Hunt Morgan
        • Cornelia Clapp
        • Ernest Everett Just
        • Ethel Browne Harvey
        • Albert Szent-Györgyi
        • Jean Clark Dan
        • Katsuma Dan
        • "Last One to Go"
        • Shinya Inoué
        • Polarized Light Microscopy
        • Seeing Spindles
        • Video Microscopy
        • Advertisements from the Collecting Net
        • Exhibit Videos
    • Alumni
      • Alumni - Overview
        • Class Photos - Overview
          • Biology of Aging Class Photos - Overview
          • Biology of Aging 2020s
          • Gene Regulatory Networks for Development Class Photos - Overview
          • Gene Regulatory Networks for Development 2000s
          • Gene Regulatory Networks for Development 2010s
          • Biology of Parasitism Class Photos - Overview
          • Biology of Parasitism 1980s
          • Biology of Parasitism 1990s
          • Biology of Parasitism 2000s
          • Biology of Parasitism 2010s
          • Biology of Parasitism 2020s
          • Brains, Minds and Machines Class Photos - Overview
          • Brains, Minds and Machines 2010s
          • Brains, Minds and Machines 2020s
          • Embryology Class Photos - Overview
          • Embryology 1880s
          • Embryology 1940s
          • Embryology 1950s
          • Embryology 1960s
          • Embryology 1970s
          • Embryology 1980s
          • Embryology 1990s
          • Embryology 2000s
          • Embryology 2010s
          • Embryology 2020s
          • Endocrine-Disrupting Chemicals: Hazards and Opportunities Class Photos - Overview
          • Endocrine-Disrupting Chemicals: Hazards and Opportunities 2020s
          • Frontiers in Reproduction Class Photos - Overview
          • Frontiers in Reproduction 1990s
          • Frontiers in Reproduction 2000s
          • Frontiers in Reproduction 2010s
          • Frontiers in Reproduction 2020s
          • Frontiers in Stem Cells and Regeneration Class Photos - Overview
          • Frontiers in Stem Cells and Regeneration 2010s
          • Frontiers in Stem Cells and Regeneration 2020s
          • History of Biology Class Photos - Overview
          • History of Biology 1980s
          • History of Biology 1990s
          • History of Biology 2000s
          • History of Biology 2010s
          • History of Biology 2020s
          • Marine Ecology Class Photos - Overview
          • Marine Ecology 1960s
          • Marine Ecology 1970s
          • Marine Ecology 1980s
          • Marine Ecology 1990s
          • Medical Informatics Class Photos - Overview
          • Medical Informatics 1990s
          • Medical Informatics 2000s
          • Microbial Diversity Class Photos - Overview
          • Microbial Diversity 1970s
          • Microbial Diversity 1980s
          • Microbial Diversity 1990s
          • Microbial Diversity 2000s
          • Microbial Diversity 2010s
          • Molecular and Cell Biology of Symbiosis Class Photos - Overview
          • Molecular and Cell Biology of Symbiosis 2020s
          • Molecular Evolution Class Photos - Overview
          • Molecular Evolution 2000s
          • Molecular Evolution 2010s
          • Molecular Mycology Class Photos - Overview
          • Molecular Mycology 1990s
          • Molecular Mycology 2000s
          • Molecular Mycology 2010s
          • Molecular Mycology 2020s
          • Neural Systems & Behavior Class Photos - Overview
          • Neural Systems & Behavior 1970s
          • Neural Systems & Behavior 1980s
          • Neural Systems & Behavior 1990s
          • Neural Systems & Behavior 2000s
          • Neural Systems & Behavior 2010s
          • Neural Systems & Behavior 2020s
          • Neurobiology Class Photos - Overview
          • Neurobiology 1970s
          • Neurobiology 1980s
          • Neurobiology 1990s
          • Neurobiology 2000s
          • Neurobiology 2010s
          • Neurobiology 2020s
          • Physiology Class Photos - Overview
          • Physiology 1890s
          • Physiology 1950s
          • Physiology 1960s
          • Physiology 1970s
          • Physiology 1980s
          • Physiology 1990s
          • Physiology 2000s
          • Physiology 2010s
          • Physiology 2020s
          • Semester in Environmental Science Class Photos - Overview
          • SES 2000s
          • SES 2010s
          • SES 2020s
          • Summer Program in Neuroscience, Excellence and Success Class Photos - Overview
          • SPINES 2020s
          • Zebrafish Class Photos - Overview
          • Zebrafish 2020s
        • Embryology 130th Anniversary Celebration - Overview
        • Symposium Schedule
        • In-Person Registrants
        • Impact Statements
      • SES 25th Anniversary Celebration and SBD Welcome Symposium
        • Neurobiology 50th Anniversary Symposium - Overview
        • Symposium Schedule
        • In-Person Registrants
        • Microbial Diversity 50th Anniversary Symposium - Overview
        • Symposium Schedule
        • In-Person Registrants
      • Contact Alumni Staff
    • Campus Facilities & Resources
      • Campus Facilities & Resources - Overview
        • Housing - Overview
        • Apply for Housing
        • Campus Info for Guests
        • Reservation Process
        • General Information and Rates
        • Housing Services
        • Housing Policies
        • Housing Buildings
          • Housing Buildings - Overview
          • Brick Dorms and Apartments
          • Memorial Circle Cottages
          • Devil's Lane Cottages
          • David House
          • Drew House
          • Smith Cottage
          • 11 North Street
          • Ebert Hall
          • Swope Building
          • Crane House
          • Veeder House
        • Local Rentals
        • Contact Housing Staff
        • Dining - Overview
        • Dining Venues
        • Dietary Accommodations
        • Swope Café Weekly Specials
        • Dining Programs
        • MBL Dining Hall Meals
        • Sodexo Sustainability @ MBL
        • Contact Dining Staff
      • Campus Maps
        • Parking, Transportation and Grounds - Overview
        • Day Visitor Parking Request
          • MBL Shuttle Service - Overview
          • MBL Policy For Shuttle Van Use
        • Contact Transportation & Grounds Staff
        • Information Technology - Overview
        • Multimedia Services
        • Telecommunications
      • Shipping and Receiving
      • Security and Campus Emergencies
      • Supplier User Guide
      • Summer Camp & Children's Programs
      • Bicycle Rentals
      • Boating Club
      • Campus Accessibility
      • Single User Restrooms
    • Local Area Information
      • Local Area Information - Overview
      • Travel Information
      • Local Hotels
      • Sculptures on the MBL Campus
    • Sustainability
      • Sustainability - Overview
      • Recycling
      • Composting
      • Electric Vehicles
      • Nitrogen and Carbon Footprint Tracking
      • Water Kiosk and Bottle Refill Stations
    • Friends of the MBL/Get Involved
    • Production Studio
    • Campus Health
    • Groundbreaking Research at the MBL
    • Research Centers
      • Research Centers - Overview
        • Ecosystems Center - Overview
        • About The Ecosystems Center
          • Ecosystems Center 50th Anniversary Celebration - Overview
          • 50th Anniversary Program
        • Ecosystems Education Info
        • Employment/Internships
          • Research Projects - Overview
          • Long-Term Ecological Research (LTER)
          • MBL Greenhouse
            • Oceanic Flux Program (OFP) - Overview
            • OFP Collaborations & Outreach
              • OFP Collaborations & Outreach - Overview
              • Agreement for the Distribution of Oceanic Flux Program Sediment Trap Material and Data
            • OFP Bibliography
            • Microscopic Photos
            • OFP Cruise Photos
            • OFP Staff
            • Contact Us
          • Terrestrial Isotope Biogeochemistry
            • Ecosystem Modeling - Overview
            • Frontiers Model
            • Hydra Model
              • General Ecosystem Model (GEM) - Overview
              • GEM Download Page
              • Multiple Element Limitation (MEL) - Overview
              • MBL MEL IV: Eight resources
              • MBL MEL V: Annual model
              • MEL Download Page
              • MEL II: Two elemental resources
            • Modelshell
            • Non-Equilibrium, Stable-Isotope Simulator (NESIS)
            • Terrestrial Ecosystem Model (TEM)
            • Tropical Forest Regrowth (TFR)
        • Center Reports
          • Stable Isotope Laboratory - Overview
          • Our Equipment
          • Sample Requirements
          • Services & Charges
          • Shipping & Turnaround
        • Media from the Lab
        • Ecosystems Publications
        • Ecosystems Center Faculty & Staff
          • Ecosystems Center Faculty & Staff - Overview
          • Paul A. Steudler
        • Eugene Bell Center - Overview
        • About the Bell Center
        • Bell Center Faculty and Staff
          • Bell Center Faculty and Staff - Overview
            • Hanlon Lab - Overview
            • About Hanlon Lab
            • Hanlon Book and Book Chapters
            • Hanlon Lab Course Material
            • Hanlon Lab in the News
            • Hanlon Photography
            • Hanlon Publications
            • Hanlon Research Areas
            • Hanlon Videos
            • Horb Lab - Overview
            • Horb Research
          • Kate MacCord
            • Oldenbourg Lab - Overview
            • Oldenbourg Gallery
            • Oldenbourg Publications
            • Rosenthal Lab - Overview
            • High-level RNA editing in cephalopods
            • Marine Model Organism Development
            • Rosenthal Lab Members
            • Site-directed RNA editing
            • Structural and functional connectivity of squid chromatophores
            • Shribak Lab - Overview
            • Shribak Gallery
        • Featured Projects
          • Featured Projects - Overview
            • Seeing Cells Exhibit - Overview
            • Seeing Cells in Life
            • Theory of Cells as Basic Units of Life
            • Protoplasm as the Living Stuff
            • Cell Specialization
            • From Fertilization to Organisms
            • Presenting and Representing "The" Cell
            • Experimental Manipulations
            • Resolving the Inside of Cells
            • Inside Living Cells
            • Seeing How Cell Parts Move
            • Seeing Cell Aggregates
            • Designing Synthetic Cells
            • Imagining Cells
            • Seeing Cells Videos
            • Tropical Seaweed Farm Project - Overview
            • Our Research
            • Tropical Seaweed News
              • Tropical Seaweed News - Overview
              • Loretta Roberson is Funded to Implement Seaweed Farms for Biomass Production
              • Meet our new project coordinator, Dr. Gretchen Grebe!
              • Members of the ANCIENT MARINER consortium successfully install 24,500 ft of line with young kelp sporophytes in Kodiak, AK
              • Our first research farm is in the water!
              • Part 1: Damisela’s Concept, Construction, and Cruise to Puerto Rico
              • The Nature Conservancy and the Belize Women’s Seaweed Farmers Association featured on The Fish Site
            • Tropical Seaweed Field Sites
            • Project Team
            • Photo Gallery
            • Press
          • Bell Center Publications - Overview
          • Bell Center Publications 2024
          • Bell Center Publications 2023
          • Bell Center Publications 2022
          • Bell Center Publications 2021
        • Contact Bell Center
        • Josephine Bay Paul Center - Overview
        • About the JPBC
          • Bay Paul Publications - Overview
          • JBPC Publications 2024
          • JBPC Publications 2023
          • JBPC Publications 2022
          • JBPC Publications 2021
          • JBPC Publications: 2011 - 2020
          • JBPC Publications: 2000 - 2010
          • JBPC Publications: 1996 - 1999
          • The Keck Facility - Overview
          • Illumina Marker Gene and Shotgun Sequencing
          • Illumina Sequencing Requirements
        • Mobile Genetic Element Cluster
          • 2024 CGUE Conference on Unicellular Eukaryotes - Overview
          • CGUE Registration and Travel Information
          • CGUE Speakers
          • CGUE Early Career Events and More
          • CGUE Program
        • Bay Paul Center Faculty and Staff
        • Whitman Center - Overview
        • About the Whitman Center
        • Registration and Housing
          • Whitman Directory - Overview
          • 2024 Whitman Fellows
          • 2024 Whitman Scientists
          • Pre-arrival, Arrival & Departure - Overview
          • Getting Started
          • Labs, Equipment, and Facilities
          • Responsible Conduct
          • Shut Down and Winter Storage
    • Faculty Directory
    • Whitman Directory
    • Researchers and Support Staff
    • Strategic Initiatives
      • Strategic Initiatives - Overview
      • Imaging Innovation Initiative
      • New Research Organisms
    • Research Organisms
      • Research Organisms - Overview
      • African Clawed Frog
      • Atlantic Horseshoe Crab
      • Axolotl
      • Bat Sea Star
      • California Two-Spot Octopus
      • Comb Jelly
      • Common Cuttlefish
      • Flamboyant Cuttlefish
      • Green Algae
      • Hummingbird Bobtail Squid
      • Lesser Pacific Striped Octopus
      • Little Skate
      • Longfin Squid
      • Northern Star Coral
      • Parhyale
      • Rotifer
      • Sea Lamprey
      • Sea Robin
      • Striped Bass
    • MBL/UChicago PhD Graduate Program
    • Scientific Publications
      • Scientific Publications - Overview
      • 2024 Scientific Publications
      • 2023 Scientific Publications
    • Who is at the MBL
      • Who is at the MBL - Overview
      • Adjunct Scientists
        • MBL Postdoctoral Association - Overview
          • 2025 Postdoctoral Symposium - Overview
          • 2025 Postdoctoral Symposium Schedule
        • PDA Events
        • PDA Volunteer Opportunities
    • MBLWHOI Library
    • Resources & Research Facilities
      • Resources & Research Facilities - Overview
        • Imaging Resources - Overview
        • Microscopes in Development
        • Central Microscopy Facility - Overview
        • CMF Equipment
        • Scheduling Time and Trainings
        • MLSC Imaging Community
        • Contact CMF
        • Laboratory Operations - Overview
        • Reserve Equipment
          • Reserve Equipment - Overview
          • Balances
          • Microscopy
        • General Use Rooms
        • Submersible Heaters
        • Stockroom Information
        • Marine Resources Center - Overview
        • About the MRC
        • MRC Catalog
        • MRC Services
        • Small Boat Program
        • Research at the MRC
          • Cephalopods - Overview
          • Cephalopod Breeding Center
          • Order Organisms & Services - Overview
          • Ordering Vertebrates
          • Tank Space Form
        • Gemma Tracker
        • Contact MRC Staff
      • The Microbiome Center
        • Zebrafish Facility - Overview
        • Zebrafish Use
        • The National Xenopus Resource - Overview
        • About the NXR
        • Ordering and Pricing
        • Protocols
          • Protocols - Overview
          • NXR Cryosperm IVF (Straws)
          • NXR Cryosperm IVF (Tubes)
          • NXR Sperm Cryopreservation Protocol (X. laevis)
        • 3D Printing Files
        • Mutant Frog Stocks
        • Transgenic Tropicalis
        • Transgenic Laevis
          • Transgenic Laevis - Overview
          • Xla.Tg (CMV:Brainbow-1.0L) NXR_0002
          • Xla.Tg(LCMV:eYFPeCFP)Ryff NXR_0003
        • Wild Type Frog Stocks
        • Research Facility Resource
        • Workshops and Meetings
          • Workshops and Meetings - Overview
          • XRET Program
        • Contact The NXR
        • Diving at MBL - Overview
        • Dive Manual
      • 10X Genomics Chromium Controller
      • Live Organism Use Form
    • Research Administration
      • Research Administration - Overview
      • Contact Research Staff
        • Office of Sponsored Programs - Overview
          • Sponsored Programs - Overview
          • Types of Agreements & Roles
          • Gifts vs. Grants
          • Uniform Guidance Information - Overview
          • Federal Agency Guidelines and Procurement
          • Subawards
          • Costs and Fees
          • Admin Salaries and Conferences
          • PI Absence from Project
          • Closeout
          • Grant & Contract Administration - Overview
          • Conflict of Interest and PHS (Public Health Science) Sponsored Specific Requirements
          • Effort Reporting: Semiannual Payroll Verification
          • Federal Policies & OSP Procedures
          • Funding & Sponsor Information
          • NIH and NSF Registration Requests
          • NSF Off-Campus/Off-Site Research (SAIF)
        • OSP Forms
        • All OSP News
          • All OSP News - Overview
          • Announcing a Simplified Review Framework for NIH Research Project Grant Applications
          • NIH Grants Family-Friendly Initiatives
          • NIH Simplified Peer Review Framework
          • NIH: Preparing for 2025 Application and Review Changes – Tips and Resources
          • NIH’s Adoption of Common Forms for Biographical Sketch and Current and Pending (Other) Support by May 25, 2025
          • Single Audit Requirements for Foreign Recipients and Subrecipients of NIH Grants
          • Updated Guidance on Salary Limitation for Grants and Cooperative Agreements
          • eRA Information: New Data Management and Sharing (DMS) Policy Questions in RPPR as of October 1, 2024, and Other DMS Changes
          • Behavioral Codes of Conduct for NIH Award Recipients
          • FORMS-H: Instructions, Forms, and a Handy Checklist
          • FY 2022 Updated Guidance: Requirement for Instruction in the Responsible Conduct of Research
          • How to Create a Grants.gov Account and Link to a Login.gov Account
          • How to Get Help When Using the Grants.gov System
          • Issuance of the NSF Proposal and Award Policies and Procedures Guide (PAPPG) (NSF 24-1)
          • NIH Application Instruction Updates – Data Management and Sharing (DMS) Costs
          • NIH Guidance on Marking Changes in Resubmission Applications
          • NSF Award Terms and Conditions Revised to Implement Build America, Buy America
          • NSF Pre-award and Post-award Disclosures Relating to the Biographical Sketch and Current and Pending Support
          • NSF Project Report Enhancements for Datasets and Research Materials
          • NSF-PAR System Updates Including Two Additional Product Types
          • Overview of NIH Grant Application and Review Changes for Due Dates on or after January 25, 2025
          • Reporting Data Management and Sharing (DMS) Plan Activities in the Research Performance Progress Report (RPPR)
          • Revised NSF Biographical Sketch and Current and Pending (Other) Support Formats - Now Available in SciENcv and on the NSF Website
          • Revision of NSF Award Terms and Conditions Applicable On or After May 20, 2024
          • Salary Cap Guidance Effective January 1, 2024
          • Updated Information about the Data Management and Sharing Plan from the NSF Directorate of Biological Sciences
          • eRA Update: Refreshed eRA Website Launched
        • Contact OSP
        • For Foreign Nationals - Overview
          • Exchange Visitors Program - Overview
          • J-1 Visa Instructions
          • English Proficiency Policy for International Participants
        • Resources
          • Tax Information for Recipients of Scholarships, Fellowships, Stipends and Honoraria - Overview
          • Applying for SSN or Individual Taxpayer ID
          • Foreign National Information System (FNIS)
          • Visas, Employment Restrictions, and Payments
        • Types of Visas
        • Contact Foreign Nationals Staff
        • Compliance and Ethics - Overview
          • IACUC - Overview
          • IACUC Charter
          • Reporting Animal Care Concerns
          • Committee and Regulations
          • Animal Use Guidelines and Purchasing
        • Independent Review Board
        • Intellectual Property
        • Policies
        • Resources
        • Training
        • Contact HR Director
    • MBL Toolbox
    • Research Registration Forms
    • Funding Opportunities
      • Funding Opportunities - Overview
        • Whitman Center Fellowships - Overview
        • Whitman Center Fellowships Supporting Funds
        • Whitman Center Research Award Recipients
        • Whitman Center Research Awards FAQ
      • Grass Foundation
    • Health & Safety
      • Health & Safety - Overview
        • Biological Safety - Overview
        • Biosafety Committee
        • IBC Application
        • IBC Charter
        • Chemical Safety - Overview
        • Acutely Hazardous Chemicals
        • Chemical Inventory Instructions
        • Chemical Storage and Segregation
        • Peroxide Forming Chemicals (PFC) Guidelines
        • Safety Data Sheet (SDS) Information
      • DEA Controlled Substances
        • Occupational Health and Safety - Overview
        • Electrical Safety
        • Fall Protection Program
        • Fire Safety Policy
        • Forklift Safety
        • Hearing Conservation
        • Ladder Safety
        • Lockout/Tagout Program
        • Machine Guarding
        • Occupational Health Program for Animal Handlers
        • Occupational Health and Safety Program
        • Workstation Ergonomics
        • Radiation and Laser Safety - Overview
        • Radiation Safety Committee Charter
        • Radiation Safety Committee
      • Research Safety Questionnaire Form
      • Waste Management
      • Contact EHS Staff
    • Educational Opportunities
    • Advanced Research Training Courses
      • Advanced Research Training Courses - Overview
      • Course Offerings
        • Course Offerings - Overview
          • Analytical and Quantitative Light Microscopy - Overview
          • Course Faculty and Lecturers
          • Financial and Course Support
          • 2025 AQLM Schedule
          • Biology of Aging - Overview
          • Course Faculty and Lecturers
          • Financial and Course Support
          • 2023 Biology of Aging Schedule
          • Biology of Parasitism: Modern Approaches - Overview
          • Course Faculty and Lecturers
          • Financial and Course Support
          • 2024 BoP Schedule
          • Student Testimonials
          • Blout Lectureship
          • Englund Lectureship
          • Biology of the Inner Ear: Experimental and Analytical Approaches - Overview
          • Course Faculty and Lecturers
          • Financial and Course Support
          • BIE Schedule
          • BIE Protocols
          • Brains, Minds and Machines - Overview
          • Course Faculty and Lecturers
          • Financial and Course Support
          • 2024 BMM Schedule
          • AI@MBL: Machine Learning for Microscopy Image Analysis - Overview
          • Course Faculty and Lecturers
          • Financial and Course Support
          • 2025 AI@MBL Schedule
          • Embryology: Concepts and Techniques in Modern Developmental Biology - Overview
          • Course Faculty and Lecturers
          • Financial and Course Support
          • 2024 Embryology Schedule
          • Course Structure and Modules
          • Course Objectives
          • Course History
          • Named Lectureships
          • Student Testimonials
          • Embryology Publications
          • Endocrine-Disrupting Chemicals: Hazards and Opportunities - Overview
          • Course Faculty and Lecturers
          • Financial and Course Support
          • 2024 ECHO Schedule
          • Frontiers in Reproduction - Overview
          • Course Faculty and Lecturers
          • Financial and Course Support
          • 2025 FIR Schedule
          • Annual FIR Symposium
          • Course History
          • Student Testimonials
          • Awards
          • Frontiers in Stem Cells & Regeneration - Overview
          • Course Faculty and Lecturers
          • 2024 SCARE Schedule
          • Financial and Course Support
          • Gene Regulatory Networks for Development - Overview
          • Course Faculty and Lecturers
          • Financial and Course Support
          • 2025 GERN Schedule
          • Methods in Computational Neuroscience - Overview
          • Course Faculty and Lecturers
          • Financial and Course Support
          • 2024 MCN Schedule
          • Course Materials
          • Lecture Videos
          • Microbial Diversity - Overview
          • Course Faculty and Lecturers
          • Financial and Course Support
          • 2024 Microbial Diversity Schedule
          • Student Testimonials
            • Publications - Overview
            • Microbial Diversity Publications 1996-2019
          • Molecular Mycology: Current Approaches to Fungal Pathogenesis - Overview
          • Course Faculty and Lecturers
          • Financial and Course Support
          • 2024 MOMY Schedule
          • The Deborah Hogan and Andrew Alspaugh Endowed Lecture
          • The Jack Edwards and Aaron Mitchell Endowed Lecture
          • Molecular and Cell Biology of Symbiosis - Overview
          • Financial and Course Support
          • 2023 Molecular and Cell Biology of Symbiosis Schedule
          • Neural Systems & Behavior - Overview
          • Course Faculty and Lecturers
          • Financial and Course Support
          • Course Structure and Modules
          • 2025 NS&B Calendar
          • 2024 NS&B Students
          • NS&B Endowed Lectures
          • Publications
          • Video Lectures
          • Video Tutorials
          • Course History
          • Student Testimonials
          • Neurobiology: Mechanisms & Advanced Approaches - Overview
          • Course Faculty and Lecturers
          • Financial and Course Support
          • 2024 Neurobiology Schedule
          • Publications
          • Student Testimonials
          • Optical Microscopy & Imaging in the Biomedical Sciences - Overview
          • Course Faculty and Lecturers
          • Financial and Course Support
          • 2024 OMIBS Schedule
          • Physiology: Modern Cell Biology Using Microscopic, Biochemical and Computational Approaches - Overview
          • Course Faculty and Lecturers
          • Financial and Course Support
          • About Physiology
          • 2025 Physiology Schedule
          • Student Testimonials
          • Strategies and Techniques for Analyzing Microbial Population Structures - Overview
          • Course Faculty and Lecturers
          • Financial and Course Support
          • 2024 STAMPS Schedule
          • Summer Program in Neuroscience, Excellence and Success - Overview
          • Course Faculty and Lecturers
          • Financial and Course Support
          • 2024 SPINES Schedule
          • Visual Neuroscience - Overview
          • Course Faculty and Lecturers
          • Financial and Course Support
          • Workshop on Molecular Evolution - Overview
          • 2025 Molecular Evolution Schedule
          • Course Faculty and Lecturers
          • Financial and Course Support
          • Zebrafish Development and Genetics - Overview
          • Course Faculty and Lecturers
          • Financial and Course Support
          • 2024 Zebrafish Schedule
          • Student Awards Presentation
      • Faculty Opportunities in MBL Courses
      • Before You Apply
      • Apply
      • Courses Calendar
      • Lecture Schedules
        • Info for Students - Overview
        • Immigration and Tax Guidance
      • Building Innovation: A faculty enhancement program
        • MBL Alumni ROCS - Overview
        • 2023-24 Alumni ROCS Awarded Projects
        • 2022-23 Alumni ROCS Awarded Projects
        • 2021-2022 Alumni ROCS Awarded Projects
        • Info for Course Faculty and Staff - Overview
        • Info for Foreign National Course Faculty
        • Education Travel Information - Overview
        • Travel Policies
    • MBL Light Microscopy Bootcamp
    • MBL/UChicago Graduate Research Fellowship Program
    • Undergraduate Programs
      • Undergraduate Programs - Overview
        • Semester in Environmental Science - Overview
          • SES Courses - Overview
          • SES Faculty
          • 2024 SES Calendar
          • 2024 SES Syllabus
          • Distinguished Scientist Seminar Series
          • SES Research Symposium
        • SES Application Instructions
        • SES Tuition and Aid
        • SES FAQ
        • SES Program Credits
        • Participating Colleges
        • Student Life
          • SES Alumni - Overview
          • SES Students & Projects
          • SES Alumni List by College
          • Participation in Conferences
            • SES 25th Anniversary Celebration and SBD Welcome Symposium - Overview
            • SES/SBD Event Gallery
        • SES Multimedia
        • Support SES
        • SES Contacts
        • Semester in Biological Discovery (SBD) - Overview
          • 2025 SBD Course Offerings - Overview
            • Art Course: On Images - Overview
            • Art Course: On Images Schedule
            • Biodiversity: Exploring the Marine Diversity of Woods Hole Using Molecular Tools - Overview
            • Biodiversity: Exploring the Marine Diversity of Woods Hole Using Molecular Tools Schedule
            • Meet the Faculty: Andrew Gillis
            • Dynamic Camouflage: Behavior, Visual Perception and Neural Skin Patterning in Cephalopods - Overview
            • Dynamic Camouflage: Behavior, Visual Perception and Neural Skin Patterning in Cephalopods Schedule
            • Meet the Faculty: Roger Hanlon
            • Embryology - Overview
            • Embryology Schedule
            • Fundamentals of Synapses - Overview
            • Fundamentals of Synapses Schedule
          • Imaging for Biological Research
            • Introduction to Parasitology - Overview
            • Introduction to Parasitology Schedule
          • Science Writing
            • Stem Cells and Regeneration: From Aquatic Research Organisms to Mammals - Overview
            • Stem Cells and Regeneration Schedule
            • Meet the Faculty: Karen Echeverri
        • SBD Application Instructions
        • SBD Tuition and Aid
        • SBD FAQ
        • SBD Program Credits
        • Student Life
        • SBD Contacts
      • UChicago Undergraduate Programs
    • High School Science Discovery Program
      • High School Science Discovery Program - Overview
      • Frequently Asked Questions
      • Participating Schools
      • Sample Course Schedule
      • Student Experiences
    • Internships
      • Internships - Overview
        • NSF REU Biological Discovery in Woods Hole - Overview
        • NSF REU Program Details
        • NSF REU Program FAQ
        • NSF REU Abstracts and Videos
          • NSF REU Abstracts and Videos - Overview
          • 2021 Abstracts
          • 2019 Abstracts
          • 2018 Abstracts
          • 2017 Abstracts
      • NSF REU – Beaufort Lagoon Ecosystems LTER
      • REU – Oceanic Flux Program
      • NSF REU – Plum Island LTER
        • Jeff Metcalf Summer for Undergraduate Research Fellows (SURF) - Overview
        • Metcalf Mentors
        • 2019 Metcalf Abstracts
        • 2018 Metcalf Abstracts
        • 2017 Metcalf Abstracts
        • 2016 Metcalf Abstracts
        • Dartmouth College E.E. Just Internship - Overview
        • Dartmouth College E.E. Just Internship Mentors
      • The Blue Economy Internship Program (BEIP)
      • Marine Aquarist and Research Internship
      • Ecosystems Center Paid Internship
    • Logan Science Journalism Program
      • Logan Science Journalism Program - Overview
      • 2025 SJP Fellows
      • Apply for SJP
      • SJP Eligibility
        • Biomedical Course - Overview
        • 2025 Schedule
        • Environmental Course - Overview
        • 2025 Schedule
        • Alumni and Testimonials - Overview
        • 2024 SJP Fellows
        • 2023 SJP Fellows
        • 2022 SJP Fellows
        • 2021 SJP Fellows
        • Faculty and Staff - Overview
        • Emeritus Directors
      • Funding and Support
      • Contact SJP Staff
    • Comparative Developmental Biology
      • Comparative Developmental Biology - Overview
      • CDB Schedule
    • Student Accessibility Services
      • Student Accessibility Services - Overview
      • Accommodation Request Form
    • Scientific Vendor Partnerships
    • MBL Toolbox
    • Education Registration Forms
    • Contact Education Staff
    • What's Happening
    • Falmouth Forum
      • Falmouth Forum - Overview
      • 2023-24 Falmouth Forum
      • 2022-23 Falmouth Forum
      • 2021-22 Falmouth Forum
      • 2020-21 Falmouth Forum
      • 2019-20 Falmouth Forum
      • 2018-19 Falmouth Forum
      • 2017-18 Falmouth Forum
      • 2016-17 Falmouth Forum
      • 2015-16 Falmouth Forum
      • 2014-15 Falmouth Forum
      • Falmouth Forum Archives
    • Friday Evening Lectures
      • Friday Evening Lectures - Overview
      • 2023 Friday Evening Lectures
      • 2022 Friday Evening Lectures
      • 2021 Friday Evening Lectures
      • 2020 Friday Evening Lectures
      • 2019 Friday Evening Lectures
      • 2018 Friday Evening Lectures
      • 2017 Friday Evening Lectures
      • 2016 Friday Evening Lectures
      • 2015 Friday Evening Lectures
    • MBL Video Center
    • Public Events & Lectures
      • Public Events & Lectures - Overview
      • SES Distinguished Scientist Seminar Series
      • Science Before Supper
    • For Scientists & Students
      • For Scientists & Students - Overview
      • Endowed Lectures
      • Flash Talk Presentations
      • MBL Virtual Programs (MVP)
      • Monday Night Neuroscience Seminars
      • Tuesday Evening Seminar Series on Cell Division and the Cytoskeleton
      • Wednesday Evening Neural Systems & Behavior Seminars
      • Thursday Evening NeuroImaging Seminars
    • Conferences
      • Conferences - Overview
      • General Information & FAQs
      • Conference Policies
        • For Conference Organizers - Overview
          • Meeting Spaces and Other Facilities - Overview
          • Auditoriums
          • Classrooms and Conference Space
          • Multi-Function and Event Space
          • Setup Options
        • Conference Rates
        • Housing and Meals
        • Conference Add-ons
      • Upcoming Conferences at the MBL
      • Contact Conference Staff
    • MBL Virtual Backgrounds
    • All MBL News
    • Newsletter
    • Recent Grants
      • Recent Grants - Overview
      • Grant Awards 2022-23
    • MBL Accolades
    • MBL Publications
      • MBL Publications - Overview
      • Annual Reports
      • Catalyst Magazine
    • Media Information
    • Social Media Information
    • Science Communication Resources for Scientists
    • Obituaries
      • Obituaries - Overview
      • Akemi Shimomura
      • Alan Fein
      • Alfred L. Goldberg
      • Anatol Eberhard
      • Annette Wilbois Coleman
      • Anton Post
      • Beth Liles
      • Captain Bill Klimm
      • Cecily C. Selby
      • Charles Kurt Fuglister
      • Cheryl Ann Palm
      • David Remsen
      • Donald A. Abt
      • Donald Price
      • Earl H. Weidner
      • Eduardo Macagno
      • Ehud (Udi) Kaplan
      • Everett Mendelsohn
      • Frank Everett Sylvia
      • Garland Edward Allen III
      • George M. Woodwell
      • Hans Laufer
      • Harish C. Pant
      • Harold Gainer
      • Jane Florence Fessenden
      • Jelle Atema
      • Jianwu (Jim) Tang
      • John J. Lee
      • John M. Kingsbury
      • Jonathan Budinoff Wittenberg 
      • Joseph G. Gall
      • Lawrence Baruch Cohen
      • Maria Poravas
      • Mark C. Alliegro
      • Michael J. Zigmond
      • Michael John Greenberg
      • Michael Patrick Sheetz
      • Michael V.L. Bennett
      • Norman Bernstein
      • Raymond Andrew Sjodin
      • Richard R. Fay
      • Robert Haselkorn
      • Robert J. Zimmer
      • Susan Joslin
      • Thomas S. Reese
      • William T. Speck
      • Irwin W. Sherman
      • Edward O. Wilson
      • Jon Henderson
      • Kenyon S. Tweedell
      • Roger W. Hobbs, Jr.
      • Ernest Couch
      • Cynthia C. MacDonald
      • Richard E. Kendall
      • Louise Luckenbill
      • Leonard Laster
      • Joan Marie Edstrom
      • Joe L. Martinez
      • Alan Grant Lunn
      • Jeanne S. Poindexter
      • Prosser Gifford
      • James G. Townsel
      • Merle Mizell
      • Gerard L. Swope
      • Robert A. Prendergast
      • Robert F. Illgen
      • Dorothy "Dixie" Berthel
      • Frank M. Child
      • Frank Press
      • Michael Toner
      • Sir Hans Kornberg
      • Kensal E. van Holde
      • Max M. Burger
      • Shinya Inoué
      • Signhild Tamm
      • Kurt J. Isselbacher
      • Paul Greengard
      • Gerald Weissmann
      • Sydney Brenner
      • Charles M. Rosenthal
      • Leonard Warren
      • John Wilson Moore
      • David C. Gadsby
      • Arthur Pardee
      • Alfred M. Zeien
      • Melvin Spiegel
      • Paul T. Englund
      • Margaret Armstrong
      • Seymour Cohen
      • Laszlo Lorand
      • Osamu Shimomura
      • Michael Edward McClure
      • David Sheprow
      • Napoleon “Mac” McCall, Jr.
      • Raymond Lipicky
      • Shinpei Ohki
      • Jane C. Kaltenbach-Townsend
      • Morris J. Karnovsky
      • Brian Kaminer
      • John E. Lisman
      • A. Kristine Johnson
      • Eloise E. Clark
      • Lewis P. Rowland
      • Christopher M. Weld
      • Margaret J. Clowes
      • Gwen P. Szent-Györgyi
      • William D. Hummon
      • Jewel Plummer Cobb
Back to Physiology Class Photos
Physiology 1890s
  • Home
  • About The MBL
  • Alumni
  • Class Photos
  • Physiology Class Photos

Physiology 1890s

physiology 1894
1894
physiology 1899
1899
The University of Chicago Marine Biological Laboratory
MBL In Your Inbox

Sign up to receive “The Collecting Net,” our bi-weekly newsletter.

Sign up for the Collecting Net
MBL Toolbox

Quickly access resources and logistical information for students, instructors and researchers.

  • Prepare Your Lab
  • Housing Information
  • Imaging Resources
  • Order an Organism
  • What's Happening at the MBL?
  • Who Is At The MBL?
  • Access the MBL Toolbox
Connect
  • Facebook
  • Twitter
  • Instagram
  • Vimeo
  • Youtube
  • LinkedIn
Main navigation
    • About the MBL
    • Leadership
    • Advisory Bodies
    • MBL Community
    • University of Chicago
    • Financial Statements
    • Careers & Internships
    • History & Archives
    • Alumni
    • Campus Facilities & Resources
    • Local Area Information
    • Sustainability
    • Friends of the MBL/Get Involved
    • Production Studio
    • Campus Health
    • Groundbreaking Research at the MBL
    • Research Centers
    • Faculty Directory
    • Whitman Directory
    • Researchers and Support Staff
    • Strategic Initiatives
    • Research Organisms
    • MBL/UChicago PhD Graduate Program
    • Scientific Publications
    • Who is at the MBL
    • MBLWHOI Library
    • Resources & Research Facilities
    • Research Administration
    • MBL Toolbox
    • Research Registration Forms
    • Funding Opportunities
    • Health & Safety
    • Educational Opportunities
    • Advanced Research Training Courses
    • MBL Light Microscopy Bootcamp
    • MBL/UChicago Graduate Research Fellowship Program
    • Undergraduate Programs
    • High School Science Discovery Program
    • Internships
    • Logan Science Journalism Program
    • Comparative Developmental Biology
    • Student Accessibility Services
    • Scientific Vendor Partnerships
    • MBL Toolbox
    • Education Registration Forms
    • Contact Education Staff
    • What's Happening
    • Falmouth Forum
    • Friday Evening Lectures
    • MBL Video Center
    • Public Events & Lectures
    • For Scientists & Students
    • Conferences
    • MBL Virtual Backgrounds
    • All MBL News
    • Newsletter
    • Recent Grants
    • MBL Accolades
    • MBL Publications
    • Media Information
    • Social Media Information
    • Science Communication Resources for Scientists
    • Obituaries
  • MBL Gift Shop
    • MBL Intranet
    • Stay For The Season
    • Visit The MBL

© 1996-2025, The Marine Biological Laboratory, MARINE BIOLOGICAL LABORATORY, MBL, and the 1888 logo are registered trademarks and service marks of The Marine Biological Laboratory.

Subfooter
  • Privacy Policy
  • Accessibility